evolution of greek vase design peaked in athens during the fifth century bc, a period of enhancement during which figure painting took on over more natural qualities.

위에서 진하게 한 부분 해석 부탁드립니다.
특히 took on over more 때문에 더 어렵게 느껴지는데 저것은 어떻게 해결해야 하나요?